[More Letters of Charles Darwin by Charles Darwin]@TWC D-Link bookMore Letters of Charles Darwin CHAPTER 1 110/193
Waterhouse (who knows the group well, and who has published on fewness of species in aberrant genera) has given me a list of 62 aberrant genera, and these have on an average 7.6 species; and if one single genus be removed (and which I cannot yet believe ought to be considered aberrant), then the 61 aberrant genera would have only 4.91 species on an average.
I tested these results in another way.
I found in Schoenherr 9 families, including only 11 genera, and these genera (9 of which were in Waterhouse's list) I found included only 3.36 species on an average. This last result led me to Lindley's "Vegetable Kingdom," in which I found (excluding thallogens and acrogens) that the genera include each 10.46 species (how near by chance to the Curculionidae), and I find 21 orders including single genera, and these 21 genera have on average 7.95 species; but if Lindley is right that Erythroxylon (with its 75 species) ought to be amongst the Malpighiads, then the average would be only 4.6 per genus. But here comes, as it appears to me, an odd thing (I hope I shall not quite weary you out).
There are 29 other orders, each with 2 genera, and these 58 genera have on an average 15.07 species: this great number being owing to the 10 genera in the Smilaceae, Salicaceae (with 220 species), Begoniaceae, Balsaminaceae, Grossulariaceae, without which the remaining 48 genera have on an average only 5.91 species. This case of the orders with only 2 genera, the genera notwithstanding having 15.07 species each, seems to me very perplexing and upsets, almost, the conclusion deducible from the orders with single genera. I have gone higher, and tested the alliances with 1, 2, and 3 orders; and in these cases I find both the genera few in each alliance, and the species, less than the average of the whole kingdom, in each genus. All this has amused me, but I daresay you will have a good sneer at me, and tell me to stick to my barnacles.
By the way, you agree with me that sometimes one gets despondent--for instance, when theory and facts will not harmonise; but what appears to me even worse, and makes me despair, is, when I see from the same great class of facts, men like Barrande deduce conclusions, such as his "Colonies" (41/3.
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