[On the Origin of Species by Charles Darwin]@TWC D-Link book
On the Origin of Species

CHAPTER V
35/48

We may, I think, confidently come to this conclusion, because, as we have seen, these coloured marks are eminently liable to appear in the crossed offspring of two distinct and differently coloured breeds; and in this case there is nothing in the external conditions of life to cause the reappearance of the slaty-blue, with the several marks, beyond the influence of the mere act of crossing on the laws of inheritance.
No doubt it is a very surprising fact that characters should reappear after having been lost for many, probably for hundreds of generations.
But when a breed has been crossed only once by some other breed, the offspring occasionally show for many generations a tendency to revert in character to the foreign breed--some say, for a dozen or even a score of generations.

After twelve generations, the proportion of blood, to use a common expression, from one ancestor, is only 1 in 2048; and yet, as we see, it is generally believed that a tendency to reversion is retained by this remnant of foreign blood.

In a breed which has not been crossed, but in which BOTH parents have lost some character which their progenitor possessed, the tendency, whether strong or weak, to reproduce the lost character might, as was formerly remarked, for all that we can see to the contrary, be transmitted for almost any number of generations.

When a character which has been lost in a breed, reappears after a great number of generations, the most probable hypothesis is, not that one individual suddenly takes after an ancestor removed by some hundred generations, but that in each successive generation the character in question has been lying latent, and at last, under unknown favourable conditions, is developed.

With the barb-pigeon, for instance, which very rarely produces a blue bird, it is probable that there is a latent tendency in each generation to produce blue plumage.


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