Some authors believe that long-continued domestication eliminates this strong tendency to sterility in species.

From the history of the dog, and of some other domestic animals, this conclusion is probably quite correct, if applied to species closely related to each other.

But to extend it so far as to suppose that species, aboriginally as distinct as carriers, tumblers, pouters, and fantails now are, should yield offspring perfectly fertile, inter se, seems to me rash in the extreme.
From these several reasons, namely, the improbability of man having formerly made seven or eight supposed species of pigeons to breed freely under domestication--these supposed species being quite unknown in a wild state, and their not having become anywhere feral--these species presenting certain very abnormal characters, as compared with all other Columbidae, though so like the rock-pigeon in most other respects--the occasional reappearance of the blue colour and various black marks in all the breeds, both when kept pure and when crossed--and lastly, the mongrel offspring being perfectly fertile--from these several reasons, taken together, we may safely conclude that all our domestic breeds are descended from the rock-pigeon or Columba livia with its geographical sub-species.
In favour of this view, I may add, firstly, that the wild C.livia has been found capable of domestication in Europe and in India; and that it agrees in habits and in a great number of points of structure with all the domestic breeds.

Secondly, that although an English carrier or a short-faced tumbler differs immensely in certain characters from the rock-pigeon, yet that by comparing the several sub-breeds of these two races, more especially those brought from distant countries, we can make, between them and the rock-pigeon, an almost perfect series; so we can in some other cases, but not with all the breeds.

Thirdly, those characters which are mainly distinctive of each breed are in each eminently variable, for instance, the wattle and length of beak of the carrier, the shortness of that of the tumbler, and the number of tail-feathers in the fantail; and the explanation of this fact will be obvious when we treat of selection.