[Darwinism (1889) by Alfred Russel Wallace]@TWC D-Link bookDarwinism (1889) CHAPTER VI 21/40
If less size of body would be beneficial, then, as half the variations in size are above and half below the mean or existing standard of the species, there would be ample beneficial variations; if a darker colour or a longer beak or wing were required, there are always a considerable number of individuals darker and lighter in colour than the average, with longer or with shorter beaks and wings, and thus the beneficial variation must always be present.
And so with every other part, organ, function, or habit; because, as variation, so far as we know, is and always must be in the two directions of excess and defect in relation to the mean amount, whichever kind of variation is wanted is always present in some degree, and thus the difficulty as to "beneficial" variations occurring, as if they were a special and rare class, falls to the ground.
No doubt some organs may vary in three or perhaps more directions, as in the length, breadth, thickness, or curvature of the bill.
But these may be taken as separate variations, each of which again occurs as "more" or "less"; and thus the "right" or "beneficial" or "useful" variation must always be present so long as any variation at all occurs; and it has not yet been proved that in any large or dominant species, or in any part, organ, or faculty of such species, there is no variation.
And even were such a case found it would prove nothing, so long as in numerous other species variation was shown to exist; because we know that great numbers of species and groups throughout all geological time have died out, leaving no descendants; and the obvious and sufficient explanation of this fact is, that they did _not_ vary enough at the time when variation was required to bring them into harmony with changed conditions.
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