I have altered the fifth edition of the _Origin_ so as to confine my remarks to adaptive changes of structure, _but I am convinced, from the light gained during even the last few years, that very many structures which now appear to us useless, will hereafter be proved to be useful, and will therefore come within the range of natural selection_.

Nevertheless I did not formerly consider sufficiently the existence of structures which, _as far as we can at present judge_, are neither beneficial nor injurious; and this I believe to be one of the greatest oversights as yet detected in my work." Now it is to be remarked that neither in these passages nor in any of the other less distinct expressions of opinion on this question, does Darwin ever admit that "specific characters"-- that is, the particular characters which serve to distinguish one species from another--are ever useless, much less that "a large proportion of them" are so, as Mr.Romanes makes him "freely acknowledge." On the other hand, in the passage which I have italicised he strongly expresses his view that much of what we suppose to be useless is due to our ignorance; and as I hold myself that, as regards many of the supposed useless characters, this is the true explanation, it may be well to give a brief sketch of the progress of knowledge in transferring characters from the one category to the other.
We have only to go back a single generation, and not even the most acute botanist could have suggested a reasonable use, for each species of plant, of the infinitely varied forms, sizes, and colours of the flowers, the shapes and arrangement of the leaves, and the numerous other external characters of the whole plant.

But since Mr.Darwin showed that plants gained both in vigour and in fertility by being crossed with other individuals of the same species, and that this crossing was usually effected by insects which, in search of nectar or pollen, carried the pollen from one plant to the flowers of another plant, almost every detail is found to have a purpose and a use.

The shape, the size, and the colour of the petals, even the streaks and spots with which they are adorned, the position in which they stand, the movements of the stamens and pistil at various times, especially at the period of, and just after, fertilisation, have been proved to be strictly adaptive in so many cases that botanists now believe that all the external characters of flowers either are or have been of use to the species.
It has also been shown, by Kerner and other botanists, that another set of characteristics have relation to the prevention of ants, slugs, and other animals from reaching the flowers, because these creatures would devour or injure them without effecting fertilisation.

The spines, hairs, or sticky glands on the stem or flower-stalk, the curious hairs or processes shutting up the flower, or sometimes even the extreme smoothness and polish of the outside of the petals so that few insects can hang to the part, have been shown to be related to the possible intrusion of these "unbidden guests."[42] And, still more recently, attempts have been made by Grant Allen and Sir John Lubbock to account for the innumerable forms, textures, and groupings of leaves, by their relation to the needs of the plants themselves; and there can be little doubt that these attempts will be ultimately successful.