He was contemporary with the southern elephant, the rhinoceros leptorhinus, the great hippopotamus, perhaps even in the miocene contemporary with the mastodon.
At the close of the Tertiary period, from causes not yet determined, the Northern Hemisphere underwent a great depression of temperature.

From a torrid it passed to a glacial condition.

After a period of prodigious length, the temperature again rose, and the glaciers that had so extensively covered the surface receded.

Once more there was a decline in the heat, and the glaciers again advanced, but this time not so far as formerly.

This ushered in the Quaternary period, during which very slowly the temperature came to its present degree.